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t is the
incipient plant on the specific morphological tendency" does not sound
to my ears like good mother-English--it wants translating. Here and
there you might, I think, have condensed some sentences. I go on the
plan of thinking every single word which can be omitted without actual
loss of sense as a decided gain. Now perhaps you will think me a
meddling intruder: anyhow, it is the advice of an old hackneyed
writer who sincerely wishes you well. Your remark on the two sexes
counteracting variability in product of the one is new to me. (151/2.
Scott (op. cit., page 214): "The reproductive organs of phoenogams,
as is well-known, are always products of two morphologically distinct
organs, the stamens producing the pollen, the carpels producing the
ovules...The embryo being in this case the modified resultant of two
originally distinct organs, there will necessarily be a greater tendency
to efface any individual peculiarities of these than would have been
the case had the embryo been the product of a single organ." A different
idea seems to have occurred to Mr. Darwin, for in an undated letter
to Scott he wrote: "I hardly know what to say on your view of male and
female organs and variability. I must think more over it. But I was
amused by finding the other day in my portfolio devoted to bud-variation
a slip of paper dated June, 1860, with some such words as these, 'May
not permanence of grafted buds be due to the two sexual elements
derived from different parts not having come into play?' I had utterly
forgotten, when I read your paper that any analogous notion had ever
passed through my mind--nor can I now remember, but the slip shows me
that it had." It is interesting that Huxley also came to a conclusion
differing from Scott's; and, curiously enough, Darwin confused the two
views, for he wrote to Scott (December 19th): "By an odd chance, reading
last night some short lectures just published by Prof. Huxley, I find
your observation, independently arrived at by him, on the confluence of
the two sexes causing variability." Professor Huxley's remarks are in
his "Lectures to Working Men on our Knowledge, etc." No. 4, page 90:
"And, indeed, I think that a certain amount of variation from the
primitive stock is the necessary result of the method of sexual
propagation itself; for inasmuch as the thing propagated proceeds from
two organisms of different sexes and different makes and temperaments,
and, as the offspring is to be either of one sex or the other, it is
quite clear that it cannot be an exact diagonal of the two, or it would
be of no sex at all; it cannot be an exact intermediate form between
that of each of its parents--it must deviate to one side or the other.")
But I cannot avoid thinking that there is something unknown and deeper
in seminal generation. Reflect on the long succession of embryological
changes in every animal. Does a bud ever produce cotyledons or embryonic
leaves? I have been much interested by your remark on inheritance at
corresponding ages; I hope you will, as you say, continue to attend to
this. Is it true that female Primula plants always produce females
by parthenogenesis? (151/3. It seems probable that Darwin here means
vegetative reproduction.) If you can answer this I should be glad;
it bears on my Primula work. I thought on the subject, but gave
up investigating what had been observed, because the female bee by
parthenogenesis produces males alone. Your paper has told me much that
in my ignorance was quite new to me. Thanks about P. scotica. If any
important criticisms are made on the Primula to the Botanical Society,
I should be glad to hear them. If you think fit, you may state that I
repeated the crossing experiments on P. sinensis and cowslip with the
same result this spring as last year--indeed, with rather more marked
difference in fertility of the two crosses. In fact, had I then proved
the Linum case, I would not have wasted time in repetition. I am
determined I will at once publish on Linum...

I was right to be cautious in supposing you in error about Siphocampylus
(no flowers were enclosed). I hope that you will make out whether the
pistil presents two definite lengths; I shall be astounded if it does. I
do not fully understand your objections to Natural Selection; if I do,
I presume they would apply with full force to, for instance, birds.
Reflect on modification of Arab-Turk horse into our English racehorse. I
have had the satisfaction to tell my publisher to send my "Journal" and
"Origin" to your address. I suspect, with your fertile mind, you
will find it far better to experiment on your own choice; but if, on
reflection, you would like to try some which interest me, I should be
truly delighted, and in this case would write in some detail. If
you have the means to repeat Gartner's experiments on variations of
Verbascum or on maize (see the "Origin"), such experiments would be
pre-eminently important. I could never get variations of Verbascum.
I could suggest an experiment on potatoes analogous with the case of
Passiflora; even the case of Passiflora, often as it has been repeated,
might be with advantage repeated. I have worked like a slave (having
counted about nine thousand seeds) on Melastoma, on the meaning of the
two sets of very different stamens, and as yet have been shamefully
beaten, and I now cry for aid. I could suggest what I believe a
very good scheme (at least, Dr. Hooker thought so) for systematic
degeneration of culinary plants, and so find out their origin; but this
would be laborious and the work of years.

LETTER 152. TO J.D. HOOKER. Down, 12th [December, 1862].

My good old Friend--

How kind you have been to give me so much of your time! Your letter
is of real use, and has been and shall be well considered. I am much
pleased to find that we do not differ as much as I feared. I begin my
book with saying that my chief object is to show the inordinate scale
of variation; I have especially studied all sorts of variations of the
individual. On crossing I cannot change; the more I think, the more
reason I have to believe that my conclusion would be agreed to by
all practised breeders. I also greatly doubt about variability and
domestication being at all necessarily correlative, but I have touched
on this in "Origin." Plants being identical under very different
conditions has always seemed to me a very heavy argument against what
I call direct action. I think perhaps I will take the case of 1,000
pigeons (152/1. See Letter 146.) to sum up my volume; I will not discuss
other points, but, as I have said, I shall recur to your letter. But
I must just say that if sterility be allowed to come into play, if
long-beaked be in the least degree sterile with short-beaked, my whole
case is altered. By the way, my notions on hybridity are becoming
considerably altered by my dimorphic work. I am now strongly inclined to
believe that sterility is at first a selected quality to keep incipient
species distinct. If you ha

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