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ncipal of a College! I proceeded to discuss his
Geology with the effrontery of a novice; and, thank God, I urged the
very argument of your letter about evidence of subsidence--viz., not all
submerged at once, and glacial action being subaerial and not oceanic.
Your letter hence was a relief, for I felt I was hardly strong enough to
have launched out as I did to a professed geologist.

(144/1. [On the subject of the above letter, see one of earlier date by
Sir J.D. Hooker (November 2nd, 1862) given in the present work (Letter
354) with Darwin's reply (Letter 355).])

LETTER 145. TO HUGH FALCONER. Down, November 14th [1862].

I have read your paper (145/1. "On the disputed Affinity of the
Mammalian Genus Plagiaulax, from the Purbeck beds."--"Quart. Journ.
Geol. Soc." Volume XVIII., page 348, 1862.) with extreme interest, and I
thank you for sending it, though I should certainly have carefully
read it, or anything with your name, in the Journal. It seems to me a
masterpiece of close reasoning: although, of course, not a judge of
such subjects, I cannot feel any doubt that it is conclusive. Will Owen
answer you? I expect that from his arrogant view of his own position he
will not answer. Your paper is dreadfully severe on him, but perfectly
courteous, and polished as the finest dagger. How kind you are towards
me: your first sentence (145/2. "One of the most accurate observers and
original thinkers of our time has discoursed with emphatic eloquence on
the Imperfection of the Geological Record.") has pleased me more than
perhaps it ought to do, if I had any modesty in my composition. By the
way, after reading the first whole paragraph, I re-read it, not for
matter, but for style; and then it suddenly occurred to me that a
certain man once said to me, when I urged him to publish some of his
miscellaneous wealth of knowledge, "Oh, he could not write,--he hated
it," etc. You false man, never say that to me again. Your incidental
remark on the remarkable specialisation of Plagiaulax (145/3. "If
Plagiaulax be regarded through the medium of the view advocated with
such power by Darwin, through what a number of intermediate forms must
not the genus have passed before it attained the specialised condition
in which the fossils come before us!") (which has stuck in my gizzard
ever since I read your first paper) as bearing on the number of
preceding forms, is quite new to me, and, of course, is in accordance to
my notions a most impressive argument. I was also glad to be reminded of
teeth of camel and tarsal bones. (145/4. Op. cit. page 353. A reference
to Cuvier's instance "of the secret relation between the upper
canine-shaped incisors of the camel and the bones of the tarsus.")
Descent from an intermediate form, Ahem!

Well, all I can say is that I have not been for a long time more
interested with a paper than with yours. It gives me a demoniacal
chuckle to think of Owen's pleasant countenance when he reads it.

I have not been in London since the end of September; when I do come I
will beat up your quarters if I possibly can; but I do not know what
has come over me. I am worse than ever in bearing any excitement. Even
talking of an evening for less than two hours has twice recently brought
on such violent vomiting and trembling that I dread coming up to London.
I hear that you came out strong at Cambridge (145/5. Prof. Owen, in a
communication to the British Association at Cambridge (1862) "On a tooth
of Mastodon from the Tertiary marls, near Shanghai," brought forward
the case of the Australian Mastodon as a proof of the remarkable
geographical distribution of the Proboscidia. In a subsequent discussion
he frankly abandoned it, in consequence of the doubts then urged
regarding its authenticity. (See footnote, page 101, in Falconer's paper
"On the American Fossil Elephant," "Nat. Hist. Review," 1863.)), and am
heartily glad you attacked the Australian Mastodon. I never did or could
believe in him. I wish you would read my little Primula paper in the
"Linnean Journal," Volume VI. Botany (No. 22), page 77 (I have no copy
which I can spare), as I think there is a good chance that you may have
observed similar cases. This is my real hobby-horse at present. I have
re-tested this summer the functional difference of the two forms in
Primula, and find all strictly accurate. If you should know of any cases
analogous, pray inform me. Farewell, my good and kind friend.

LETTER 146. TO J.D. HOOKER.

(146/1. The following letter is interesting in connection with a letter
addressed to Sir J.D. Hooker, March 26th, 1862, No. 136, where the
value of Natural Selection is stated more strongly by Sir Joseph than by
Darwin. It is unfortunate that Sir Joseph's letter, to which this is a
reply, has not been found.)

Down, November 20th [1862].

Your last letter has interested me to an extraordinary degree, and your
truly parsonic advice, "some other wise and discreet person," etc.,
etc., amused us not a little. I will put a concrete case to show what
I think A. Gray believes about crossing and what I believe. If 1,000
pigeons were bred together in a cage for 10,000 years their number
not being allowed to increase by chance killing, then from mutual
intercrossing no varieties would arise; but, if each pigeon were a
self-fertilising hermaphrodite, a multitude of varieties would
arise. This, I believe, is the common effect of crossing, viz., the
obliteration of incipient varieties. I do not deny that when two marked
varieties have been produced, their crossing will produce a third
or more intermediate varieties. Possibly, or probably, with domestic
varieties, with a strong tendency to vary, the act of crossing tends
to give rise to new characters; and thus a third or more races, not
strictly intermediate, may be produced. But there is heavy evidence
against new characters arising from crossing wild forms; only
intermediate races are then produced. Now, do you agree thus far? if
not, it is no use arguing; we must come to swearing, and I am convinced
I can swear harder than you, therefore I am right. Q.E.D.

If the number of 1,000 pigeons were prevented increasing not by chance
killing, but by, say, all the shorter-beaked birds being killed, then
the WHOLE body would come to have longer beaks. Do you agree?

Thirdly, if 1,000 pigeons were kept in a hot country, and another
1,000 in a cold country, and fed on different food, and confined in
different-size aviary, and kept constant in number by chance killing,
then I should expect as rather probable that after 10,000 years the two
bodies would differ slightly in size, colour, and perhaps other trifling
characters; this I should call the direct action of physical conditions.
By this action I wish to imply that the innate vital forces are somehow
led to act rather differently in the two cases, just as heat will
allow or cause two elements to combine, which otherwise would not have
combined. I should be especially obliged if you would tell me what you
think on 

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