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nder the present state of
science, believe in the creation of a few aboriginal forms, or of some
one form of life. In the sixth edition, probably with a view to the
umbilicus, he writes (page 423): "Undoubtedly some of these same
questions," etc., etc. From notes in Mr. Darwin's copy of the second
edition it is clear that the change in the third edition was chiefly
due to Harvey's letter. See Letter 115.) about my question (page 483,
"Origin") about creation of eggs or young, etc., (but not about mammals
with the mark of the umbilical cord), yet I still have an illogical sort
of feeling that there is less difficulty in imagining the creation of an
asexual cell, increasing by simple division.

Page 5 of your letter: I agree to every word about the antiquity of the
world, and never saw the case put by any one more strongly or more ably.
It makes, however, no more impression on me as an objection than does
the astronomer when he puts on a few hundred million miles to the
distance of the fixed stars. To compare very small things with great,
Lingula, etc., remaining nearly unaltered from the Silurian epoch to
the present day, is like the dovecote pigeons still being identical with
wild Rock-pigeons, whereas its "fancy" offspring have been immensely
modified, and are still being modified, by means of artificial
selection.

You put the difficulty of the first modification of the first protozoon
admirably. I assure you that immediately after the first edition was
published this occurred to me, and I thought of inserting it in the
second edition. I did not, because we know not in the least what the
first germ of life was, nor have we any fact at all to guide us in our
speculations on the kind of change which its offspring underwent. I
dissent quite from what you say of the myriads of years it would take to
people the world with such imagined protozoon. In how very short a time
Ehrenberg calculated that a single infusorium might make a cube of rock!
A single cube on geometrical progression would make the solid globe
in (I suppose) under a century. From what little I know, I cannot help
thinking that you underrate the effects of the physical conditions of
life on these low organisms. But I fully admit that I can give no sort
of answer to your objections; yet I must add that it would be marvellous
if any man ever could, assuming for the moment that my theory is true.
You beg the question, I think, in saying that Protococcus would be
doomed to eternal similarity. Nor can you know that the first germ
resembled a Protococcus or any other now living form.

Page 12 of your letter: There is nothing in my theory necessitating in
each case progression of organisation, though Natural Selection tends in
this line, and has generally thus acted. An animal, if it become
fitted by selection to live the life, for instance, of a parasite, will
generally become degraded. I have much regretted that I did not make
this part of the subject clearer. I left out this and many other
subjects, which I now see ought to have been introduced. I have inserted
a discussion on this subject in the foreign editions. (110/7. In the
third Edition a discussion on this point is added in Chapter IV.) In
no case will any organic being tend to retrograde, unless such
retrogradation be an advantage to its varying offspring; and it is
difficult to see how going back to the structure of the unknown supposed
original protozoon could ever be an advantage.

Page 13 of your letter: I have been more glad to read your discussion
on "dominant" forms than any part of your letter. (110/8. Harvey
writes: "Viewing organic nature in its widest aspect, I think it is
unquestionable that the truly dominant races are not those of high, but
those of low organisation"; and goes on to quote the potato disease,
etc. In the third edition of the "Origin," page 56, a discussion is
introduced defining the author's use of the term "dominant.") I can now
see that I have not been cautious enough in confining my definition
and meaning. I cannot say that you have altered my views. If Botrytis
[Phytophthora] had exterminated the wild potato, a low form would have
conquered a high; but I cannot remember that I have ever said (I am
sure I never thought) that a low form would never conquer a high. I have
expressly alluded to parasites half exterminating game-animals, and
to the struggle for life being sometimes between forms as different as
possible: for instance, between grasshoppers and herbivorous quadrupeds.
Under the many conditions of life which this world affords, any group
which is numerous in individuals and species and is widely distributed,
may properly be called dominant. I never dreamed of considering that
any one group, under all conditions and throughout the world, would be
predominant. How could vertebrata be predominant under the conditions
of life in which parasitic worms live? What good would their perfected
senses and their intellect serve under such conditions? When I have
spoken of dominant forms, it has been in relation to the multiplication
of new specific forms, and the dominance of any one species has been
relative generally to other members of the same group, or at least to
beings exposed to similar conditions and coming into competition. But I
daresay that I have not in the "Origin" made myself clear, and space has
rendered it impossible. But I thank you most sincerely for your valuable
remarks, though I do not agree with them.

About sudden jumps: I have no objection to them--they would aid me in
some cases. All I can say is, that I went into the subject, and found
no evidence to make me believe in jumps; and a good deal pointing in the
other direction. You will find it difficult (page 14 of your letter) to
make a marked line of separation between fertile and infertile crosses.
I do not see how the apparently sudden change (for the suddenness of
change in a chrysalis is of course largely only apparent) in larvae
during their development throws any light on the subject.

I wish I could have made this letter better worth sending to you. I have
had it copied to save you at least the intolerable trouble of reading
my bad handwriting. Again I thank you for your great liberality and
kindness in sending me your criticisms, and I heartily wish we were
a little nearer in accord; but we must remain content to be as wide
asunder as the poles, but without, thank God, any malice or other
ill-feeling.

LETTER 111. TO T.H. HUXLEY.

(111/1. Dr. Asa Gray's articles in the "Atlantic Monthly," July, August,
and October, 1860, were published in England as a pamphlet, and form
Chapter III. in his "Darwiniana" (1876). See "Life and Letters," II.,
page 338. The article referred to in the present letter is that in the
August number.)

Down, September 10th [1860].

I send by this post a review by Asa Gray, so good that I should like you
to see it; I must beg for its return. I want to ask, also, your opinion
about getting it reprinted in England. I th

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