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w groan, and think to yourself, "on what a
man have I been wasting my time and writing to." I should, five years
ago, have thought so...(13/3. On the questions here dealt with see the
interesting letter to Jenyns in the "Life and Letters," II., page 34.)

LETTER 14. TO J.D. HOOKER. [November] 1844.

...What a curious, wonderful case is that of the Lycopodium! (14/1. Sir
J.D. Hooker wrote, November 8, 1844: "I am firmly convinced (but not
enough to print it) that L. Selago varies in Van Diemen's Land into L.
varium. Two more different SPECIES (as they have hitherto been thought),
per se cannot be conceived, but nowhere else do they vary into one
another, nor does Selago vary at all in England.")...I suppose you would
hardly have expected them to be more varying than a phanerogamic plant.
I trust you will work the case out, and, even if unsupported, publish
it, for you can surely do this with due caution. I have heard of some
analogous facts, though on the smallest scale, in certain insects being
more variable in one district than in another, and I think the same
holds with some land-shells. By a strange chance I had noted to ask you
in this letter an analogous question, with respect to genera, in lieu
of individual species,--that is, whether you know of any case of a genus
with most of its species being variable (say Rubus) in one continent,
having another set of species in another continent non-variable, or not
in so marked a manner. Mr. Herbert (14/2. No doubt Dean Herbert, the
horticulturist. See "Life and Letters," I., page 343.) incidentally
mentioned in a letter to me that the heaths at the Cape of Good Hope
were very variable, whilst in Europe they are (?) not so; but then the
species here are few in comparison, so that the case, even if true, is
not a good one. In some genera of insects the variability appears to be
common in distant parts of the world. In shells, I hope hereafter to
get much light on this question through fossils. If you can help me, I
should be very much obliged: indeed, all your letters are most useful to
me.

MONDAY:--Now for your first long letter, and to me quite as interesting
as long. Several things are quite new to me in it--viz., for one, your
belief that there are more extra-tropical than intra-tropical species. I
see that my argument from the Arctic regions is false, and I should not
have tried to argue against you, had I not fancied that you thought that
equability of climate was the direct cause of the creation of a greater
or lesser number of species. I see you call our climate equable; I
should have thought it was the contrary. Anyhow, the term is vague, and
in England will depend upon whether a person compares it with the United
States or Tierra del Fuego. In my Journal (page 342) I see I state that
in South Chiloe, at a height of about 1,000 feet, the forests had a
Fuegian aspect: I distinctly recollect that at the sea-level in the
middle of Chiloe the forest had almost a tropical aspect. I should like
much to hear, if you make out, whether the N. or S. boundaries of a
plant are the most restricted; I should have expected that the S. would
be, in the temperate regions, from the number of antagonist species
being greater. N.B. Humboldt, when in London, told me of some river
(14/3. The Obi (see "Flora Antarctica," page 211, note). Hooker writes:
"Some of the most conspicuous trees attain either of its banks, but do
not cross them.") in N.E. Europe, on the opposite banks of which the
flora was, on the same soil and under same climate, widely different!

I forget (14/4. The last paragraph is published in "Life and Letters,"
II., page 29.) my last letter, but it must have been a very silly one,
as it seems I gave my notion of the number of species being in great
degree governed by the degree to which the area had been often isolated
and divided. I must have been cracked to have written it, for I have no
evidence, without a person be willing to admit all my views, and then it
does follow.

(14/5. The remainder of the foregoing letter is published in the "Life
and Letters," II., page 29. It is interesting as giving his views on
the mutability of species. Thus he wrote: "With respect to books on this
subject, I do not know any systematical ones, except Lamarck's, which is
veritable rubbish; but there are plenty, as Lyell, Pritchard, etc., on
the view of the immutability." By "Pritchard" is no doubt intended James
Cowles "Prichard," author of the "Physical History of Mankind." Prof.
Poulton has given in his paper, "A remarkable Anticipation of Modern
Views on Evolution" (14/6. "Science Progress," Volume I., April 1897,
page 278.), an interesting study of Prichard's work. He shows that
Prichard was in advance of his day in his views on the non-transmission
of acquired characters. Prof. Poulton also tries to show that Prichard
was an evolutionist. He allows that Prichard wrote with hesitation, and
that in the later editions of his book his views became weaker.
But, even with these qualifications, we think that Poulton has
unintentionally exaggerated the degree to which Prichard believed in
evolution.

One of Prichard's strongest sentences is quoted by Poulton (loc. cit.,
page 16); it occurs in the "Physical History of Mankind," Ed. 2, Volume
II., page 570:--

"Is it not probable that the varieties which spring up within the
limits of particular species are further adaptations of structure to
the circumstances under which the tribe is destined to exist? Varieties
branch out from the common form of a species, just as the forms of
species deviate from the common type of a genus. Why should the
one class of phenomena be without end or utility, a mere effect of
contingency or chance, more than the other?"

If this passage, and others similar to it, stood alone, we might agree
with Prof. Poulton; but this is impossible when we find in Volume I.
of the same edition, page 90, the following uncompromising statement of
immutability:--

"The meaning attached to the term species, in natural history, is
very simple and obvious. It includes only one circumstance--namely, an
original distinctness and constant transmission of any character. A race
of animals, or plants, marked by any peculiarities of structure which
have always been constant and undeviating, constitutes a species."

On page 91, in speaking of the idea that the species which make up a
genus may have descended from a common form, he says:--

"There must, indeed, be some principle on which the phenomena of
resemblance, as well as those of diversity, may be explained; and the
reference of several forms to a common type seems calculated to suggest
the idea of some original affinity; but, as this is merely a conjecture,
it must be kept out of sight when our inquiries respect matters of fact
only."

This view is again given in Volume II., page 569, where he asks whether
we should believe that "at the first production of a genus, when it
first grew into existence, some slight modification in th

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