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stressed
by fall of temperature, and probably by changed humidity, and changed
proportional numbers of other plants and enemies (though they might
favour some of the species), I must admit that they all would be
exterminated if productions exactly fitted, not only for the climate,
but for all the conditions of the equatorial regions during the Glacial
period existed and could everywhere have immigrated. But the productions
of the temperate regions would have probably found, under the equator,
in their new homes and soils, considerably different conditions of
humidity and periodicity, and they would have encountered a new set of
enemies (a most important consideration); for there seems good reason
to believe that animals were not able to migrate nearly to the extent to
which plants did during the Glacial period. Hence I can persuade
myself that the temperate productions would not entirely replace and
exterminate the productions of the cooled tropics, but would become
partially mingled with them.

I am far from satisfied with what I have scribbled. I conclude that
there must have been a mundane Glacial period, and that the difficulties
are much the same whether we suppose it contemporaneous over the world,
or that longitudinal belts were affected one after the other. For
Heaven's sake forgive me!

LETTER 353. TO H.W. BATES. March 26th [1861].

I have been particularly struck by your remarks on the Glacial period.
(353/1. In his "Contributions to the Insect Fauna of the Amazon Valley,"
"Trans. Entom. Soc." Volume V., page 335 (read November 24th, 1860),
Mr. Bates discusses the migration of species from the equatorial regions
after the Glacial period. He arrives at a result which, he points
out, "is highly interesting as bearing upon the question of how far
extinction is likely to have occurred in equatorial regions during
the time of the Glacial epoch."..."The result is plain, that there
has always (at least throughout immense geological epochs) been an
equatorial fauna rich in endemic species, and that extinction cannot
have prevailed to any extent within a period of time so comparatively
modern as the Glacial epoch in geology." This conclusion does not
support the view expressed in the "Origin of Species" (Edition I.,
chapter XI., page 378) that the refrigeration of the earth extended to
the equatorial regions. (Bates, loc. cit., pages 352, 353.)) You seem
to me to have put the case with admirable clearness and with crushing
force. I am quite staggered with the blow, and do not know what to
think. Of late several facts have turned up leading me to believe more
firmly that the Glacial period did affect the equatorial regions; but I
can make no answer to your argument, and am completely in a cleft
stick. By an odd chance I have only a few days ago been discussing
this subject, in relation to plants, with Dr. Hooker, who believes to a
certain extent, but strongly urged the little apparent extinction in the
equatorial regions. I stated in a letter some days ago to him that the
tropics of S. America seem to have suffered less than the Old World.
There are many perplexing points; temperate plants seem to have migrated
far more than animals. Possibly species may have been formed more
rapidly within tropics than one would have expected. I freely confess
that you have confounded me; but I cannot yet give up my belief that the
Glacial period did to certain extent affect the tropics.

LETTER 354. TO J.D. HOOKER. Down, February 25th [1862].

I have almost finished your Arctic paper, and I must tell you how I
admire it. (354/1. "Outlines of the Distribution of Arctic Plants"
[Read June 21st, 1860], "Linn. Soc. Trans." XXIII., 1862, page 251. The
author's remarks on Mr. Darwin's theories of Geographical Distribution
are given at page 255: they are written in a characteristically
generous spirit.) The subject, treated as you have treated it, is really
magnificent. Good Heaven, what labour it must have cost you! And what a
grand prospect there is for the future. I need not say how much pleased
I am at your notice of my work; for you know that I regard your opinion
more than that of all others. Such papers are the real engine to compel
people to reflect on modification of species; any one with an enquiring
mind could hardly fail to wish to consider the whole subject after
reading your paper. By Jove! you will be driven, nolens volens, to a
cooled globe. Think of your own case of Abyssinia and Fernando Po, and
South Africa, and of your Lebanon case (354/2. See "Origin," Edition
VI., page 337.); grant that there are highlands to favour migration, but
surely the lowlands must have been somewhat cooled. What a splendid new
and original evidence and case is that of Greenland: I cannot see how,
even by granting bridges of continuous land, one can understand the
existing flora. I should think from the state of Scotland and America,
and from isothermals, that during the coldest part of Glacial period,
Greenland must have been quite depopulated. Like a dog to his vomit, I
cannot help going back and leaning to accidental means of transport
by ice and currents. How curious also is the case of Iceland. What a
splendid paper you have made of the subject. When we meet I must ask you
how much you attribute richness of flora of Lapland to mere climate; it
seems to me very marvellous that this point should have been a sort of
focus of radiation; if, however, it is unnaturally rich, i.e. contains
more species than it ought to do for its latitude, in comparison with
the other Arctic regions, would it not thus falsely seem a focus of
radiation? But I shall hereafter have to go over and over again your
paper; at present I am quite muddy on the subject. How very odd, on any
view, the relation of Greenland to the mountains of E. N. America; this
looks as if there had been wholesale extinction in E. N. America. But I
must not run on. By the way, I find Link in 1820 speculated on relation
of Alpine and Arctic plants being due to former colder climate, which he
attributed to higher mountains cutting off the warm southern winds.

LETTER 355. J.D. HOOKER TO CHARLES DARWIN. Kew, November 2nd, 1862.

Did I tell you how deeply pleased I was with Gray's notice of my Arctic
essay? (355/1. "American Journal of Science and Arts," XXXIV., and in
Gray's "Scientific Papers," Volume I., page 122.) It was awfully good of
him, for I am sure he must have seen several blunders. He tells me that
Dr. Dawson (355/2. A letter (No. 144) by Sir J.D. Hooker, dated November
7th, 1862, on this subject occurs in the Evolutionary section.) is down
on me, and I have a very nice lecture on Arctic and Alpine plants from
Dr. D., with a critique on the Arctic essay--which he did not see till
afterwards. He has found some mares' nests in my essay, and one very
venial blunder in the tables--he seems to HATE Darwinism--he accuses me
of overlooking the geological facts, and dwells much on my overlooking
subsidence of temperate America during Glaci

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