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y not-trained American plants would not
succeed in our agrarian habitats.

LETTER 343. TO J.D. HOOKER.

(343/1. Mr. Darwin used the knowledge of the spread of introduced plants
in North America and Australia to throw light on the cosmic migration
of plants. Sir J.D. Hooker apparently objected that it was not fair
to argue from agrarian to other plants; he also took a view differing
slightly from that of Darwin as to climatal and other natural conditions
favouring introduced plants in Australia.)

Down, January 28th, 1859.

Thanks about glaciers. It is a pleasure and profit to me to write to
you, and as in your last you have touched on naturalised plants of
Australia, I suppose you would not dislike to hear what I can say
in answer. At least I know you would not wish me to defer to your
authority, as long as not convinced.

I quite agree to what you say about our agrarian plants being accustomed
to cultivated land, and so no fair test. Buckman has, I think, published
this notion with respect to North America. With respect to roadside
plants, I cannot feel so sure that these ought to be excluded, as
animals make roads in many wild countries. (343/2. In the account of
naturalised plants in Australia in Sir J.D. Hooker's "Introductory Essay
to the Flora of Tasmania," 1859, page cvi, many of the plants are marked
"Britain--waste places," "Europe--cornfields," etc. In the same list
the species which have also invaded North America--a large number--are
given. On the margin of Darwin's copy is scribbled in pencil: "Very
good, showing how many of the same species are naturalised in Australia
and United States, with very different climates; opposed to your
conclusion." Sir Joseph supposed that one chief cause of the intrusion
of English plants in Australia, and not vice versa, was the great
importation of European seed to Australia and the scanty return of
Australian seed.)

I have now looked and found passage in F. Muller's (343/3. Ferdinand
Muller.) letter to me, in which he says: "In the WILDERNESSES of
Australia some European perennials are "advancing in sure progress,"
"not to be arrested," etc. He gives as instances (so I suppose there
are other cases) eleven species, viz., 3. Rumex, Poterium sanguisorba,
Potentilla anserina, Medicago sativa, Taraxacum officinale, Marrubium
vulgare, Plantago lanceolata, P. major, Lolium perenne. All these are
seeding freely. Now I remember, years and years ago, your discussing
with me how curiously easily plants get naturalised on uninhabited
islands, if ships even touch there. I remember we discussed packages
being opened with old hay or straw, etc. Now think of hides and wool
(and wool exported largely over Europe), and plants introduced, and
samples of corn; and I must think that if Australia had been the old
country, and Europe had been the Botany Bay, very few, very much
fewer, Australian plants would have run wild in Europe than have now in
Australia.

The case seems to me much stronger between La Plata and Spain.

Nevertheless, I will put in my one sentence on this head, illustrating
the greater migration during Glacial period from north to south than
reversely, very humbly and cautiously. (343/4. "Origin of Species,"
Edition I., page 379. Darwin refers to the facts given by Hooker and De
Candolle showing a stronger migratory flow from north to south than in
the opposite direction. Darwin accounts for this by the northern plants
having been long subject to severe competition in their northern homes,
and having acquired a greater "dominating power" than the southern
forms. "Just in the same manner as we see at the present day that very
many European productions cover the ground in La Plata, and in a lesser
degree in Australia, and have to a certain extent beaten the natives;
whereas extremely few southern forms have become naturalised in any part
of Europe, though hides, wool, and other objects likely to carry seeds
have been largely imported during the last two or three centuries from
La Plata, and during the last thirty or forty years from Australia.')

I am very glad to hear you are making good progress with your Australian
Introduction. I am, thank God, more than half through my chapter on
geographical distribution, and have done the abstract of the Glacial
part...

LETTER 344. TO J.D. HOOKER. Down, March 30th, 1859.

Many thanks for your agreeable note. Please keep the geographical MS.
till you hear from me, for I may have to beg you to send it to Murray;
as through Lyell's intervention I hope he will publish, but he requires
first to see MS. (344/1. "The Origin of Species"; see a letter to Lyell
in "Life and Letters," II., page 151.)

I demur to what you say that we change climate of the world to account
for "migration of bugs, flies, etc." WE do nothing of the sort; for WE
rest on scored rocks, old moraines, arctic shells, and mammifers. I have
no theory whatever about cause of cold, no more than I have for cause of
elevation and subsidence; and I can see no reason why I should not use
cold, or elevation, or subsidence to explain any other phenomena, such
as distribution. I think if I had space and time I could make a pretty
good case against any great continental changes since the Glacial
epoch, and this has mainly led me to give up the Lyellian doctrine as
insufficient to explain all mutations of climate.

I was amused at the British Museum evidence. (344/2. This refers to the
letter to Murchison (Letter 65), published with the evidence of the 1858
enquiry by the Trustees of the British Museum.) I am made to give my
opinion so authoritatively on botanical matters!...

As for our belief in the origin of species making any difference in
descriptive work, I am sure it is incorrect, for I did all my barnacle
work under this point of view. Only I often groaned that I was not
allowed simply to decide whether a difference was sufficient to deserve
a name.

I am glad to hear about Huxley--a wonderful man.

LETTER 345. TO J.D. HOOKER. Wells Terrace, Ilkley, Otley, Yorkshire,
Thursday [before December 9th, 1859].

I have read your discussion (345/1. See "Introductory Essay," page
c. Darwin did not receive this work until December 23rd, so that the
reference is to proof-sheets.), as usual, with great interest. The
points are awfully intricate, almost at present beyond the confines
of knowledge. The view which I should have looked at as perhaps most
probable (though it hardly differs from yours) is that the whole world
during the Secondary ages was inhabited by marsupials, araucarias
(Mem.--Fossil wood so common of this nature in South America (345/2.
See Letter 6, Note.)), Banksia, etc.; and that these were supplanted and
exterminated in the greater area of the north, but were left alive in
the south. Whence these very ancient forms originally proceeded seems a
hopeless enquiry.

Your remarks on the passage of the northern forms southward, and of
the southern forms of no kinds passing northward, seem to me grand.
Admirable, also, are 

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