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tion, are very clearly
marked indeed.

Mentha, we have only one wild species; that has two pretty well-marked
forms, which have been taken for species; one smooth, the other hairy.

Saxifraga, gives no trouble here.

Myosotis, only one or two species here, and those very well marked.

Hieracium, few species, but pretty well marked.

Rosa, putting down a set of nominal species, leaves us four; two of them
polymorphous, but easy to distinguish...

LETTER 339. TO J.D. HOOKER. Down, [1857?]

One must judge by one's own light, however imperfect, and as I have
found no other book (339/1. A. De Candolle's "Geographie Botanique,"
1855.) so useful to me, I am bound to feel grateful: no doubt it is
in main part owing to the concentrated light of the noble art of
compilation. (339/2. See Letter 49.) I was aware that he was not the
first who had insisted on range of Monocots. (Was not R. Brown [with]
Flinders?) (339/3. M. Flinders' "Voyage to Terra Australis in 1801-3,
in H.M.S. 'Investigator'"; with "Botanical Appendix," by Robert Brown,
London, 1814.), and I fancy I only used expression "strongly insisted
on,"--but it is quite unimportant.

If you and I had time to waste, I should like to go over his [De
Candolle's] book and point out the several subjects in which I fancy he
is original. His remarks on the relations of naturalised plants will be
very useful to me; on the ranges of large families seemed to me good,
though I believe he has made a great blunder in taking families instead
of smaller groups, as I have been delighted to find in A. Gray's last
paper. But it is no use going on.

I do so wish I could understand clearly why you do not at all believe in
accidental means of dispersion of plants. The strongest argument which
I can remember at this instant is A. de C., that very widely ranging
plants are found as commonly on islands as over continents. It is really
provoking to me that the immense contrast in proportion of plants in New
Zealand and Australia seems to me a strong argument for non-continuous
land; and this does not seem to weigh in the least with you. I wish I
could put myself in your frame of mind. In Madeira I find in Wollaston's
books a parallel case with your New Zealand case--viz., the striking
absence of whole genera and orders now common in Europe, and (as I have
just been hunting out) common in Europe in Miocene periods. Of course
I can offer no explanation why this or that group is absent; but if the
means of introduction have been accidental, then one might expect odd
proportions and absences. When we meet, do try and make me see more
clearly than I do, your reasons.

LETTER 340. TO J.D. HOOKER. Down, November 14th [1858].

I am heartily glad to hear that my Lyellian notes have been of the
slightest use to you. (340/1. The Copley Medal was given to Sir Charles
Lyell in 1858. Mr. Darwin supplied Sir J.D. Hooker, who was on the
Council of the Royal Society, with notes for the reasons for the award.
See Letter 69.) I do not think the view is exaggerated...

Your letter and lists have MOST DEEPLY interested me. First for less
important point, about hermaphrodite trees. (340/2. See "Life and
Letters," II., page 89. In the "Origin," Edition I., page 100, the
author quotes Dr. Hooker to the effect that "the rule does not hold in
Australia," i.e., that trees are not more generally unisexual than other
plants. In the 6th edition, page 79, Darwin adds, "but if most of the
Australian trees are dichogamous, the same result would follow as if
they bore flowers with separated sexes.") It is enough to knock me down,
yet I can hardly think that British N. America and New Zealand should
all have been theoretically right by chance. Have you at Kew any
Eucalyptus or Australian Mimosa which sets its seeds? if so, would it
be very troublesome to observe when pollen is mature, and whether
pollen-tubes enter stigma readily immediately that pollen is mature or
some little time afterwards? though if pollen is not mature for some
little time after flower opens, the stigma might be ready first, though
according to C.C. Sprengel this is a rarer case. I wrote to Muller for
chance of his being able and willing to observe this.

Your fact of greater number of European plants (N.B.--But do you mean
greater percentage?) in Australia than in S. America is astounding and
very unpleasant to me; for from N.W. America (where nearly the
same flora exists as in Canada?) to T. del Fuego, there is far more
continuous high land than from Europe to Tasmania. There must have,
I should think, existed some curious barrier on American High-Road:
dryness of Peru, excessive damp of Panama, or some other confounded
cause, which either prevented immigration or has since destroyed them.
You say I may ask questions, and so I have on enclosed paper; but it
will of course be a very different thing whether you will think them
worth labour of answering.

May I keep the lists now returned? otherwise I will have them copied.

You said that you would give me a few cases of Australian forms
and identical species going north by Malay Archipelago mountains to
Philippines and Japan; but if these are given in your "Introduction"
this will suffice for me. (340/3. See Hooker's "Introductory Essay,"
page l.)

Your lists seem to me wonderfully interesting.

According to my theoretical notions, I am not satisfied with what you
say about local plants in S.W. corner of Australia (340/4. Sir Joseph
replied in an undated letter: "Thanks for your hint. I shall be very
cautious how I mention any connection between the varied flora and poor
soil of S.W. Australia...It is not by the way only that the species
are so numerous, but that these and the genera are so confoundedly well
marked. You have, in short, an incredible number of VERY LOCAL, WELL
MARKED genera and species crowded into that corner of Australia." See
"Introductory Essay to the Flora of Tasmania," 1859, page li.), and the
seeds not readily germinating: do be cautious on this; consider lapse
of time. It does not suit my stomach at all. It is like Wollaston's
confined land-snails in Porto Santo, and confined to same spots since
a Tertiary period, being due to their slow crawling powers; and yet we
know that other shell-snails have stocked a whole country within a very
few years with the same breeding powers, and same crawling powers,
when the conditions have been favourable to the life of the introduced
species. Hypothetically I should rather look at the case as owing
to--but as my notions are not very simple or clear, and only
hypothetical, they are not worth inflicting on you.

I had vowed not to mention my everlasting Abstract (340/5. The "Origin
of Species" was abbreviated from the MS. of an unpublished book.) to you
again, for I am sure I have bothered you far more than enough about
it; but as you allude to its previous publication I may say that I
have chapters on Instinct and Hybridism to abstract, which may take
a fortnight each; and my materials for Palaeon

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