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I have formerly discussed the notion of the world
or great belts of it having been cooler, though he at first saw great
difficulties (and difficulties there are great enough), I think is
much inclined to adopt the idea. With modification of specific forms it
explains some wondrous odd facts in distribution.

But I shall never stop if I get on this subject, on which I have been at
work, sometimes in triumph, sometimes in despair, for the last month.

LETTER 328. ASA GRAY TO CHARLES DARWIN. Received August 20th, 1856.

I enclose you a proof of the last page, that you may see what our flora
amounts to. The genera of the Cryptogams (Ferns down to Hepaticae) are
illustrated in fourteen crowded plates. So that the volume has become
rather formidable as a class-book, which it is intended for.

I have revised the last proofs to-day. The publishers will bring it out
some time in August. Meanwhile, I am going to have a little holiday,
which I have earned, little as I can spare the time for it. And my wife
and I start on Friday to visit my mother and friends in West New York,
and on our way back I will look in upon the scientific meeting at Albany
on the 20th inst., or later, just to meet some old friends there.

Why could not you come over, on the urgent invitation given to European
savans--and free passage provided back and forth in the steamers? Yet
I believe nobody is coming. Will you not come next year, if a special
invitation is sent you on the same terms?

Boott lately sent me your photograph, which (though not a very perfect
one) I am well pleased to have...

But there is another question in your last letter--one about which a
person can only give an impression--and my impression is that, speaking
of plants of a well-known flora, what we call intermediate varieties are
generally less numerous in individuals than the two states which they
connect. That this would be the case in a flora where things are put as
they naturally should be, I do not much doubt; and the wider are
your views about species (say, for instance, with Dr. Hooker's very
latitudinarian notions) the more plainly would this appear. But
practically two things stand hugely in the way of any application of the
fact or principle, if such it be. 1. Our choice of what to take as the
typical forms very often is not free. We take, e.g., for one of them the
particular form of which Linnaeus, say, happened to have a specimen sent
him, and on which [he] established the species; and I know more than one
case in which that is a rare form of a common species; the other variety
will perhaps be the opposite extreme--whether the most common or not, or
will be what L. or [illegible] described as a 2nd species. Here various
intermediate forms may be the most abundant. 2. It is just the same
thing now, in respect to specimens coming in from our new western
country. The form which first comes, and is described and named,
determines the specific character, and this long sticks as the type,
though in fact it may be far from the most common form. Yet of plants
very well known in all their aspects, I can think of several of which
we recognise two leading forms, and rarely see anything really
intermediate, such as our Mentha borealis, its hairy and its smooth
varieties.

Your former query about the variability of naturalised plants as
compared with others of same genera, I had not forgotten, but have
taken no steps to answer. I was going hereafter to take up our list of
naturalised plants and consider them--it did not fall into my plan to
do it yet. Off-hand I can only say that it does not strike me that our
introduced plants generally are more variable, nor as variable, perhaps,
as the indigenous. But this is a mere guess. When you get my sheets of
first part of article in "Silliman's Journal," remember that I shall
be most glad of free critical comments; and the earlier I get them the
greater use they will be to me...

One more favour. Do not, I pray you, speak of your letters troubling me.
I should be sorry indeed to have you stop, or write more rarely, even
though mortified to find that I can so seldom give you the information
you might reasonably expect.

LETTER 329. TO ASA GRAY. Down, August 24th [1856].

I am much obliged for your letter, which has been very interesting to
me. Your "indefinite" answers are perhaps not the least valuable part;
for Botany has been followed in so much more a philosophical spirit
than Zoology, that I scarcely ever like to trust any general remark
in Zoology without I find that botanists concur. Thus, with respect to
intermediate varieties being rare, I found it put, as I suspected, much
too strongly (without the limitations and doubts which you point out) by
a very good naturalist, Mr. Wollaston, in regard to insects; and if it
could be established as true it would, I think, be a curious point.
Your answer in regard to the introduced plants not being particularly
variable, agrees with an answer which Mr. H.C. Watson has sent me in
regard to British agrarian plants, or such (whether or no naturalised)
[as] are now found only in cultivated land. It seems to me very odd,
without any theoretical notions of any kind, that such plants should not
be variable; but the evidence seems against it.

Very sincere thanks for your kind invitation to the United States: in
truth there is nothing which I should enjoy more; but my health is not,
and will, I suppose, never be strong enough, except for the quietest
routine life in the country. I shall be particularly glad of the sheets
of your paper on geographical distribution; but it really is unlikely in
the highest degree that I could make any suggestions.

With respect to my remark that I supposed that there were but few
plants common to Europe and the United States, not ranging to the
Arctic regions; it was founded on vague grounds, and partly on range of
animals. But I took H.C. Watson's remarks (1835) and in the table at the
end I found that out of 499 plants believed to be common to the Old and
New World, only 110 did not range on either side of the Atlantic up to
the Arctic region. And on writing to Mr. Watson to ask whether he knew
of any plants not ranging northward of Britain (say 55 deg) which were
in common, he writes to me that he imagines there are very few; with Mr.
Syme's assistance he found some 20 to 25 species thus circumstanced,
but many of them, from one cause or other, he considered doubtful. As
examples, he specifies to me, with doubt, Chrysosplenium oppositifolium;
Isnardia palustris; Astragalus hypoglottis; Thlaspi alpestre; Arenaria
verna; Lythrum hyssopifolium.

I hope that you will be inclined to work out for your next paper, what
number, of your 321 in common, do not range to Arctic regions. Such
plants seem exposed to such much greater difficulties in diffusion. Very
many thanks for all your kindness and answers to my questions.

P.S.--If anything should occur to you on variability of naturalised or
agrarian plants, I hope that you 

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