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 It
is evident a genus might be widely diffused in two ways: 1st, by many
different species, each with restricted ranges; and 2nd, by many or
few species with wide ranges. Any light which you could throw on this I
should be very much obliged for. Thank you most kindly, also, for your
offer in a former letter to consider any other points; and at some
future day I shall be most grateful for a little assistance, but I will
not be unmerciful.

Swainson has remarked (and Westwood contradicted) that typical genera
have wide ranges: Waterhouse (without knowing these previous
remarkers) made to me the same observation: I feel a laudable doubt and
disinclination to believe any statement of Swainson; but now Waterhouse
remarks it, I am curious on the point. There is, however, so much vague
in the meaning of "typical forms," and no little ambiguity in the mere
assertion of "wide ranges" (for zoologists seldom go into strict and
disagreeable arithmetic, like you botanists so wisely do) that I feel
very doubtful, though some considerations tempt me to believe in this
remark. Here again, if you can throw any light, I shall be much obliged.
After your kind remarks I will not apologise for boring you with my
vague queries and remarks.

LETTER 315. TO J.D. HOOKER. Down, December 25th [1844]. Happy Christmas
to you.

(315/1. The following letter refers to notes by Sir J.D. Hooker which we
have not seen. Though we are therefore unable to make clear many points
referred to, the letter seems to us on the whole so interesting that it
is printed with the omission of only one unimportant sentence.

The subjects dealt with in the letter are those which were occupying
Hooker's attention in relation to his "Flora Antarctica" (1844).)

I must thank you once again for all your documents, which have
interested me very greatly and surprised me. I found it very difficult
to charge my head with all your tabulated results, but this I perfectly
well know is in main part due to that head not being a botanical one,
aided by the tables being in MS.; I think, however, to an ignoramus,
they might be made clearer; but pray mind, that this is very different
from saying that I think botanists ought to arrange their highest
results for non-botanists to understand easily. I will tell you how,
for my individual self, I should like to see the results worked out, and
then you can judge, whether this be advisable for the botanical world.

Looking at the globe, the Auckland and Campbell I., New Zealand, and
Van Diemen's Land so evidently are geographically related, that I should
wish, before any comparison was made with far more distant countries,
to understand their floras, in relation to each other; and the southern
ones to the northern temperate hemisphere, which I presume is to every
one an almost involuntary standard of comparison. To understand the
relation of the floras of these islands, I should like to see the group
divided into a northern and southern half, and to know how many species
exist in the latter--

1. Belonging to genera confined to Australia, Van Diemen's Land and
north New Zealand.

2. Belonging to genera found only on the mountains of Australia, Van
Diemen's Land, and north New Zealand.

3. Belonging to genera of distribution in many parts of the world (i.e.,
which tell no particular story).

4. Belonging to genera found in the northern hemisphere and not in the
tropics; or only on mountains in the tropics.

I daresay all this (as far as present materials serve) could be
extracted from your tables, as they stand; but to any one not familiar
with the names of plants, this would be difficult. I felt particularly
the want of not knowing which of the genera are found in the lowland
tropics, in understanding the relation of the Antarctic with the Arctic
floras.

If the Fuegian flora was treated in the analogous way (and this would
incidentally show how far the Cordillera are a high-road of genera), I
should then be prepared far more easily and satisfactorily to understand
the relations of Fuegia with the Auckland Islands, and consequently with
the mountains of Van Diemen's Land. Moreover, the marvellous facts
of their intimate botanical relation (between Fuegia and the Auckland
Islands, etc.) would stand out more prominently, after the Auckland
Islands had been first treated of under the purely geographical relation
of position. A triple division such as yours would lead me to suppose
that the three places were somewhat equally distant, and not so
greatly different in size: the relation of Van Diemen's Land seems so
comparatively small, and that relation being in its alpine plants, makes
me feel that it ought only to be treated of as a subdivision of the
large group, including Auckland, Campbell, New Zealand...

I think a list of the genera, common to Fuegia on the one hand and on
the other to Campbell, etc., and to the mountains of Van Diemen's Land
or New Zealand (but not found in the lowland temperate, and southern
tropical parts of South America and Australia, or New Zealand), would
prominently bring out, at the same time, the relation between these
Antarctic points one with another, and with the northern or Arctic
regions.

In Article III. is it meant to be expressed, or might it not be
understood by this article, that the similarity of the distant points
in the Antarctic regions was as close as between distant points in
the Arctic regions? I gather this is not so. You speak of the southern
points of America and Australia, etc., being "materially approximated,"
and this closer proximity being correlative with a greater similarity of
their plants: I find on the globe, that Van Diemen's Land and Fuegia are
only about one-fifth nearer than the whole distance between Port Jackson
and Concepcion in Chile; and again, that Campbell Island and Fuegia
are only one-fifth nearer than the east point of North New Zealand and
Concepcion. Now do you think in such immense distances, both over open
oceans, that one-fifth less distance, say 4,000 miles instead of 5,000,
can explain or throw much light on a material difference in the degree
of similarity in the floras of the two regions?

I trust you will work out the New Zealand flora, as you have commenced
at end of letter: is it not quite an original plan? and is it not very
surprising that New Zealand, so much nearer to Australia than South
America, should have an intermediate flora? I had fancied that nearly
all the species there were peculiar to it. I cannot but think you make
one gratuitous difficulty in ascertaining whether New Zealand ought to
be classed by itself, or with Australia or South America--namely, when
you seem (bottom of page 7 of your letter) to say that genera in
common indicate only that the external circumstances for their life are
suitable and similar. (315/2. On December 30th, 1844, Sir J.D. Hooker
replied, "Nothing was further from my intention than to have written
anything which would lead one to suppose that genera common to two
plac

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