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r of absorbing and finding a place
for new facts, and its capability of interpreting phenomena, which had
been previously looked upon as unaccountable anomalies..." Mr. Wallace
illustrates his statement that "a false theory will never stand this
test," by Edward Forbes' "polarity" speculations (see page 84 of
the present volume) and Macleay's "Circular" and "Quinarian System"
published in his "Horae Entomologicae," 1821, and developed by Swainson
in the natural history volumes of "Lardner's Cabinet Cyclopaedia."
Mr. Wallace says that a "considerable number of well-known naturalists
either spoke approvingly of it, or advocated similar principles, and for
a good many years it was decidedly in the ascendant...yet it quite died
out in a few short years, its very existence is now a matter of history,
and so rapid was its fall that...Swainson, perhaps, lived to be the last
man who believed in it. Such is the course of a false theory. That of
a true one is very different, as may be well seen by the progress of
opinion on the subject of Natural Selection."

Here, (page 3) follows a passage on the overwhelming importance of
Natural Selection, underlined with apparent approval in Mr. Darwin's
copy of the review.), in the "Westminster Review," in an article on
Protection has a good passage, contrasting the success of Natural
Selection and its growth with the comprehension of new classes of facts
(222/3. This rather obscure phrase may be rendered: "its power of growth
by the absorption of new facts."), with false theories, such as the
Quinarian Theory, and that of Polarity, by poor Forbes, both of
which were promulgated with high advantages and the first temporarily
accepted.

LETTER 223. TO G.H. LEWES.

(223/1. The following is printed from a draft letter inscribed by Mr.
Darwin "Against organs having been formed by direct action of medium in
distinct organisms. Chiefly luminous and electric organs and thorns."
The draft is carelessly written, and all but illegible.)

August 7th, 1868.

If you mean that in distinct animals, parts or organs, such for instance
as the luminous organs of insects or the electric organs of fishes, are
wholly the result of the external and internal conditions to which the
organs have been subjected, in so direct and inevitable a manner that
they could be developed whether of use or not to their possessor, I
cannot admit [your view]. I could almost as soon admit that the whole
structure of, for instance, a woodpecker, had thus originated; and
that there should be so close a relation between structure and external
circumstances which cannot directly affect the structure seems to me to
[be] inadmissible. Such organs as those above specified seem to me
much too complex and generally too well co-ordinated with the whole
organisation, for the admission that they result from conditions
independently of Natural Selection. The impression which I have taken,
studying nature, is strong, that in all cases, if we could collect all
the forms which have ever lived, we should have a close gradation from
some most simple beginning. If similar conditions sufficed, without the
aid of Natural Selection, to give similar parts or organs, independently
of blood relationship, I doubt much whether we should have that striking
harmony between the affinities, embryological development, geographical
distribution, and geological succession of all allied organisms. We
should be much more puzzled than we now are how to class, in a natural
method, many forms. It is puzzling enough to distinguish between
resemblance due to descent and to adaptation; but (fortunately
for naturalists), owing to the strong power of inheritance, and to
excessively complex causes and laws of variability, when the same end
or object has been gained, somewhat different parts have generally been
modified, and modified in a different manner, so that the resemblances
due to descent and adaptation can commonly be distinguished. I should
just like to add, that we may understand each other, how I suppose the
luminous organs of insects, for instance, to have been developed; but I
depend on conjectures, for so few luminous insects exist that we have
no means of judging, by the preservation to the present day of slightly
modified forms, of the probable gradations through which the organs have
passed. Moreover, we do not know of what use these organs are. We see
that the tissues of many animals, [as] certain centipedes in England,
are liable, under unknown conditions of food, temperature, etc., to
become occasionally luminous; just like the [illegible]: such luminosity
having been advantageous to certain insects, the tissues, I suppose,
become specialised for this purpose in an intensified degree; in certain
insects in one part, in other insects in other parts of the body. Hence
I believe that if all extinct insect-forms could be collected, we should
have gradations from the Elateridae, with their highly and constantly
luminous thoraxes, and from the Lampyridae, with their highly luminous
abdomens, to some ancient insects occasionally luminous like the
centipede.

I do not know, but suppose that the microscopical structure of the
luminous organs in the most different insects is nearly the same; and I
should attribute to inheritance from a common progenitor, the similarity
of the tissues, which under similar conditions, allowed them to vary
in the same manner, and thus, through Natural Selection for the same
general purpose, to arrive at the same result. Mutatis mutandis, I
should apply the same doctrine to the electric organs of fishes; but
here I have to make, in my own mind, the violent assumption that some
ancient fish was slightly electrical without having any special organs
for the purpose. It has been stated on evidence, not trustworthy, that
certain reptiles are electrical. It is, moreover, possible that the
so-called electric organs, whilst in a condition not highly developed,
may have subserved some distinct function: at least, I think, Matteucci
could detect no pure electricity in certain fishes provided with the
proper organs. In one of your letters you alluded to nails, claws,
hoofs, etc. From their perfect coadaptation with the whole rest of the
organisation, I cannot admit that they would have been formed by the
direct action of the conditions of life. H. Spencer's view that they
were first developed from indurated skin, the result of pressure on the
extremities, seems to me probable.

In regard to thorns and spines I suppose that stunted and [illegible]
hardened processes were primarily left by the abortion of various
appendages, but I must believe that their extreme sharpness and hardness
is the result of fluctuating variability and "the survival of the
fittest." The precise form, curvature and colour of the thorns I freely
admit to be the result of the laws of growth of each particular plant,
or of their conditions, internal and external. It would be an astounding
fact if any varying plant suddenly pr

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